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Coral reefs are diverse underwater ecosystems held together by calcium carbonate structures secreted by corals. Coral reefs are built by colonies of tiny animals found in marine waters that contain few nutrients. Most coral reefs are built from stony corals, which in turn consist of polyps that cluster in groups. The polyps belong to a group of animals known as Cnidaria, which also includes sea anemones and jellyfish. Unlike sea anemones, corals secrete hard carbonate exoskeletons which support and protect the coral polyps. Most reefs grow best in warm, shallow, clear, sunny and agitated waters. Often called "rainforests of the sea", shallow coral reefs form some of the most diverse ecosystems on Earth. They occupy less than 0.1% of the world's ocean surface, about half the area of France, yet they provide a home for at least 25% of all marine species,〔〔Mulhall M (Spring 2009) (Saving rainforests of the sea: An analysis of international efforts to conserve coral reefs ) Duke Environmental Law and Policy Forum 19:321–351.〕〔(Where are Corals Found? ) NOAA. Reviewed: 13 May 2011. Retrieved: 24 March 2015.〕 including fish, mollusks, worms, crustaceans, echinoderms, sponges, tunicates and other cnidarians. Paradoxically, coral reefs flourish even though they are surrounded by ocean waters that provide few nutrients. They are most commonly found at shallow depths in tropical waters, but deep water and cold water corals also exist on smaller scales in other areas. Coral reefs deliver ecosystem services to tourism, fisheries and shoreline protection. The annual global economic value of coral reefs is estimated between US$29.8-375 billion.〔〔 However, coral reefs are fragile ecosystems, partly because they are very sensitive to water temperature. They are under threat from climate change, oceanic acidification, blast fishing, cyanide fishing for aquarium fish, sunscreen use, overuse of reef resources, and harmful land-use practices, including urban and agricultural runoff and water pollution, which can harm reefs by encouraging excess algal growth.〔(【引用サイトリンク】 title= Corals reveal impact of land use )〕 == Formation == Most of the coral reefs we can see today were formed after the last glacial period when melting ice caused the sea level to rise and flood the continental shelves. This means that most modern coral reefs are less than 10,000 years old. As communities established themselves on the shelves, the reefs grew upwards, pacing rising sea levels. Reefs that rose too slowly could become drowned reefs. They are covered by so much water that there was insufficient light. Coral reefs are found in the deep sea away from continental shelves, around oceanic islands and as atolls. The vast majority of these islands are volcanic in origin. The few exceptions have tectonic origins where plate movements have lifted the deep ocean floor on the surface. In 1842 in his first monograph, ''The Structure and Distribution of Coral Reefs'', Charles Darwin set out his theory of the formation of atoll reefs, an idea he conceived during the voyage of the ''Beagle''. He theorized uplift and subsidence of the Earth's crust under the oceans formed the atolls. Darwin’s theory sets out a sequence of three stages in atoll formation. It starts with a fringing reef forming around an extinct volcanic island as the island and ocean floor subsides. As the subsidence continues, the fringing reef becomes a barrier reef, and ultimately an atoll reef. File:Atoll forming-volcano.png|Darwin’s theory starts with a volcanic island which becomes extinct File:Atoll forming-Fringing reef.png|As the island and ocean floor subside, coral growth builds a fringing reef, often including a shallow lagoon between the land and the main reef. File:Atoll forming-Barrier reef.png|As the subsidence continues, the fringing reef becomes a larger barrier reef further from the shore with a bigger and deeper lagoon inside. File:Atoll forming-Atoll.png|Ultimately, the island sinks below the sea, and the barrier reef becomes an atoll enclosing an open lagoon. Darwin predicted that underneath each lagoon would be a bed rock base, the remains of the original volcano. Subsequent drilling proved this correct. Darwin's theory followed from his understanding that coral polyps thrive in the clean seas of the tropics where the water is agitated, but can only live within a limited depth range, starting just below low tide. Where the level of the underlying earth allows, the corals grow around the coast to form what he called fringing reefs, and can eventually grow out from the shore to become a barrier reef. Where the bottom is rising, fringing reefs can grow around the coast, but coral raised above sea level dies and becomes white limestone. If the land subsides slowly, the fringing reefs keep pace by growing upwards on a base of older, dead coral, forming a barrier reef enclosing a lagoon between the reef and the land. A barrier reef can encircle an island, and once the island sinks below sea level a roughly circular atoll of growing coral continues to keep up with the sea level, forming a central lagoon. Barrier reefs and atolls do not usually form complete circles, but are broken in places by storms. Like sea level rise, a rapidly subsiding bottom can overwhelm coral growth, killing the coral polyps and the reef, due to what is called ''coral drowning''. Corals that rely on zooxanthellae can ''drown'' when the water becomes too deep for their symbionts to adequately photosynthesize, due to decreased light exposure. The two main variables determining the geomorphology, or shape, of coral reefs are the nature of the underlying substrate on which they rest, and the history of the change in sea level relative to that substrate. The approximately 20,000-year-old Great Barrier Reef offers an example of how coral reefs formed on continental shelves. Sea level was then lower than in the 21st century. As sea level rose, the water and the corals encroached on what had been hills of the Australian coastal plain. By 13,000 years ago, sea level had risen to lower than at present, and many hills of the coastal plains had become continental islands. As the sea level rise continued, water topped most of the continental islands. The corals could then overgrow the hills, forming the present cays and reefs. Sea level on the Great Barrier Reef has not changed significantly in the last 6,000 years,〔 and the age of the modern living reef structure is estimated to be between 6,000 and 8,000 years.〔(【引用サイトリンク】title=What is the Great Barrier Reef? )〕 Although the Great Barrier Reef formed along a continental shelf, and not around a volcanic island, Darwin's principles apply. Development stopped at the barrier reef stage, since Australia is not about to submerge. It formed the world's largest barrier reef, from shore, stretching for .〔(Four Types of Coral Reef ) Microdocs, Stanford Education. Retrieved January 10, 2010.〕 Healthy tropical coral reefs grow horizontally from per year, and grow vertically anywhere from per year; however, they grow only at depths shallower than because of their need for sunlight, and cannot grow above sea level. 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Coral reef」の詳細全文を読む スポンサード リンク
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